Bot. J. Linn. Soc. 110: 95-110, 1992

Ascomata: Fruiting bodies raised, brown, crust-like, circular spots, up to 750 µm in diam., with a small central ostiole; lenticular, developing beneath the host epidermis, with overlaying clypeus and variable development of stromatic tissue laterally. Ascomata one or two per stroma, lenticular, 225-285 m m high x 510-750 m m in diam., ostiolar canal lacking periphyses.

Asci: 142-207 x 14.2-19.8 m m, 8-spored, clavate to long-cylindrical, pedunculate, fissitunicate, the ends rounded with an ocular chamber and a ring not staining blue in Melzer’s reagent.

Ascospores: 42-66 x 7.0-10.6 m m, biseriate, yelow to light-brown, yellow in mass, 7-8 septate, constricted at the septa, the 2 central cells being largest, surrounded by a gelatinous sheath. The sheath is first ellipsoidal, but once released swells and forms a distinctive shape, with a central `spine' and curved polar extrusions.